Best Planet of the Apes Villain?

I think Dr Zaius will always be mine. In my opinion at least, he's definitely the most complex and intriguing. However, the fact that I know people who aren't even POTA fans that still know the name Koba really says a lot about the strength of that villain. Proximus I feel had the potential to be the greatest, but he didn't have enough screen time in my eyes to really explore his character as much as he needed.